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15 Crikey! Lake crocodilian from Central Otago

Sauropsida, Crocodilia

The first fossil crocodilian to be described from Aotearoa New Zealand is a small dark fragment of jaw from Early Miocene lake sediments in Central Otago. The specimen was found by Mike Pole near St Bathans during a study of fossil plants from Central Otago, and was described by Ralph Molnar and Mike Pole in 1997. While the fossil has been described, the genus and species to which it is belongs have not yet been formally identified.

The image shows a bone from the lower jaw of an unidentified species of crocodilian. The ID number of the specimen is OU22228. The bone is dark grey to brown and is being held between two fingers against a black background. The figure has no scale bar but the fingers indicate that the bone is relatively small and would easily fit in the palm of the hand. The texture of the bone appears rough and uneven, with several small indentations and ridges visible on its surface.

Bone from the lower jaw of an unidentified species of crocodilian (specimen OU 22228). Image credit: RE Fordyce.

The St Bathans fossil fragment is a little less than 7 cm long and is from the right lower jaw of a small crocodile. Given the proportions of the jaw in modern crocodiles, the original lower jaw was perhaps 20 cm long and the animal probably had a skull of a similar length. A Pleistocene fossil crocodilian from Fiji Volia athollandersoni was around two to three metres long and may have been a similar size to the specimen from St Bathans.

The crocodilian lower jaw came from the Bannockburn Formation. Fossils from Bannockburn Formation include freshwater mussels (Hyridella), abundant fish remains including isolated bones and otoliths (See: Robinson’s pilchard: the importance of earbones), bones of many species of ducks, shelducks and geese, and broken eggshell. Plant fossils are abundant in places and algal stromatolites or oncolites are known (See: Small boulders of fossil bacteria). The strata were deposited in an extensive shallow “Lake Manuherikia” which occupied much of what is now Central Otago.

The animal to which the jaw fragment belonged was possibly a member of the Mekosuchinae group of crocodilians. Mekosuchines are a distinct and extinct group of crocodiles first recognised in Australia and now known from other localities including Fiji (e.g. Volia athollandersoni), New Caledonia and Vanuatu. Mekosuchines originated at least 53 million years ago during the early Eocene and lived through to the Holocene. At least one mekosuchine crocodile from Vanuatu apparently co-existed with humans only a few thousand years ago.

Mekosuchines dispersed across oceans through the southwest Pacific. Judging from the known distribution of other mekosuchines in space and time, these animals could have lived in New Zealand either intermittently or continuously; an open question in New Zealand paleontology.

The St Bathans crocodilian jaw is currently on loan to Tūhura Otago Museum in Ōtepoti Dunedin where it can be seen in the Southern Land, Southern People gallery, along with many other specimens from the University of Otago Geology Museum.

—Written by R. Ewan Fordyce, with additional contributions by Daniel B Thomas

Specimen number: OU 22228 Age: 19 to 16 million years old (early Miocene, Altonian Stage)
Locality: St Bathans, Central Otago Rock Formation: Bannockburn Formation
Collected by: M Pole
Citation: Molnar RE, Pole M. 1997. A Miocene crocodilian from New Zealand. Alcheringa 21:65–70. doi.org/10.1080/03115519708619185
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