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36 Wing-like brachiopods endemic to a future Zealandia

Brachiopoda, Spiriferinida: Rastelligera sp.

During the Triassic and Jurassic about 200 species of brachiopods were present in the seas around southern Pangaea and the newly-forming Gondwana (future Zealandia). One particularly unusual group of brachiopods known as Rastelligera is well-represented in the University of Otago Geology Museum collections. This is a genus of spiriferinid brachiopods (these have a coiled spiralium which supported the lophophore during life) that are distinguished by a very long, wing-like hingeline. Around eight species are known and all are endemic to Zealandia. The first of these species was named in 1913. Most of the others were named in 1968 by JD (Doug) Campbell, a paleontologist and botanist who was on the staff of the Department of Geology from 1959 until his death in 2001.

The figure shows two images. The upper image is one half of the natural mould in grey brown sediment made by a fossil brachiopod Rastelligera. The lower image is the other half of the natural mould, but this image includes the steinkern or internal mould resting in the mould cavity. The calcareous external shell of the brachiopod has been completely dissolved away. Both images show the detailed rib pattern that would have been on the surface of the brachiopod. The fossil brachiopod was around eight centimetres wide and four centimetres tall.

Part, counterpart and steinkern (internal mould) of a specimen of the extinct brachiopod, Rastelligera. The calcareous external shell has been completely dissolved away. Image credit: JH Robinson.

These distinctive brachiopods are found throughout a 2000 metre thick sequence of Triassic rocks in Southland and South Otago from the Hokonui Hills to Nugget Point.  They became extinct at the end of the Triassic, a time when many other brachiopods also disappeared from the fossil record.

The name Rastelligera refers to the distinctive “comb-teeth” and was coined by James Hector in 1878 (in Latin, rastellus refers to the teeth on a rake). Some of these brachiopods are also described as mucronate, an unusual word which means to end abruptly in a point. The elongate, wing-like shape may have helped to anchor the brachiopods on the seabed and the many teeth ensured the animal could securely hold its valves closed. Because the shell material has usually been dissolved away over time, the fossils are often moulds showing the interior structures. The shape and morphology of the actual shell can be recreated by using a rubber or silicone mould and then more replicas can be made.

—Written by Jeffrey H Robinson and Daphne E Lee

Specimen number: OU 19325 Age: Approximately 204 million years old (late Triassic, Otapirian stage)
Locality: Otamita Stream, Hokonui Hills, Southland Rock Formation: Murihiku Supergroup
Collected by: Student on a Geology field trip in 2001
Citation: Campbell JD. 1968 Rastelligera (Brachiopoda) of the upper Triassic of New Zealand. Transactions of the Royal Society of New Zealand 6:23–37.
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